Calosoma (Australodrepa) oceanicum Perroud & Montrouzier 1864

This species, which spread to several islands of the archipelagos surrounding Australia, and in the north of the Australian continent itself, has the upper body of a metallic colour, variable from dark purple to golden greenish, the ventral side is also metallic, but always darker. As schayeri the other Australian species, oceanicum is characterized by a depression of the whole surface on the base of the pronotum. However it can be easily idenfied because the striae of the elytra are only superficially punctured and the interstriae almost smooth, the last article of maxillary palps is distincly longer in comparison with schayeri. In males the aedeagus has the apex somewhat elongated and it has a ligula with a triangular end, finally all the articles of the anterior tarsi are hairy on underside.
Differences, albeit slight, of form and color variations, had led to distinguish various subspecies: oceanicum of Nouvelle-Calédonie; timorese of Lesser Sunda; walkeri of the northern part of Australian continent; shayeri of South eastern Moluccans.
The differences between them are not constant, as the variability of single individuals is very high. At most we can distinguish between two groups of populations: one (oceanicum) that lives in the archipelagos east and north of the Australian continent and in the north of Australian continent itself, with individuals constantly olivacea ; and another one ( klynstrai ), in the facing south eastern Moluccans islands, with individuals more slender, brightly colored, often two-tone.

Calosoma (Australodrepa) oceanicum oceanicum Perroud & Montrouzier, 1864

Calosoma oceanicum Perroud & Montrouzier, 1864: 49 (lectotype: Kanala; Muséum National d'Histoire Naturelle, Paris)
Calosoma timorense Chaudoir, 1869: 367 (type: Timor, Atapoupou; Muséum National d'Histoire Naturelle, Paris)
Calosoma walkeri Waterhouse, 1898: 98 (type: Swam river; British Museum Natural History, London)
Calosoma (Australodrepa) timorensis Lapouge, 1932: 407
Calosoma (Australodrepa) walkeri Lapouge, 1932: 408
Calosoma (Australodrepa) oceanica Lapouge, 1932: 408
Australodrepa oceanicum Jeannel, 1940: 74
Australodrepa oceanicum walkeri Jeannel, 1940: 74
Calosoma oceanicum lombocense Häckel et Al., 2005: 4 (type: Lombok; coll. Häckel, Prague)

Length 26-33 mm. The nominal form includes populations with the upper part single-colored olivacea. In some individuals from the Lesser Sunda archipelago the pronotum and the elytra have a green lateral margin (timorese); moreover specimens from Lombok and Sumbava have been described as a distinct subspecies (Calosoma oceanicum lombocense Hacke et Al 2005) based on small differences in the elytral microsculture and in the punctuation of head and pronotum.
The populations of Northern Australia (walkeri) are on average of smaller size and the color of the upper body can be, very seldom more clear, up to greenish bronze.
It is found in Nouvelle-Calédonie, including the Îles Loyauté; Lesser Sunda archipelago: Wetar, Timor, Flores, Sumbava, Lombok and in Northern Australia (West Australia, Northern Territory, northern Queensland).

Material examined:
Nouvelle-Calédonie: Nouméa; Farino
Indonesia: Lesser Sunda archipelago (Wetar, Timor, Flores)

Australia: West Australia (Broome, Milly Soak); Northern Territory (Yuendumu, 650 m.)
Notes: Nocturnal, winged, caterpillar feeder. Active individuals were captured from October to Februry during the relatively less humid and hottest period of the year, before the beginning of the raining season.

Calosoma (Australodrepa) oceanicum oceanicum
Perroud & Montrouzier, 1864
Kanala (lectotype)
(coll. Muséum National d'Histoire Naturelle, Paris)
Calosoma (Australodrepa) oceanicum oceanicum
Perroud & Montrouzier, 1864
Atapoupou, Timor (type of Calosoma timorense Chaudoir)
(coll. Muséum National d'Histoire Naturelle, Paris)

Calosoma (Australodrepa) oceanicum oceanicum
Perroud & Montrouzier, 1864
New Caledonia: Poe beach, I.2005 (coll. Pontuale)
Calosoma (Australodrepa) oceanicum oceanicum
Perroud & Montrouzier, 1864
New Caledonia: Poe beach, I.2005 (coll. Pontuale)
Calosoma (Australodrepa) oceanicum oceanicum
Perroud & Montrouzier, 1864
Indonesia: Lesser Sundas, Wetar island, North coast,
Anpala env., 14.I.2007, S. Jaki leg., (coll. Pontuale)
Calosoma (Australodrepa) oceanicum oceanicum
Perroud & Montrouzier, 1864
Indonesia: Lesser Sundas, Flores island, X.2005, (coll. Pontuale)

Calosoma (Australodrepa) oceanicum oceanicum
Perroud & Montrouzier, 1864
West Australia: Cue Milly Soak, 6.2.2003, Hovorka lg.
Calosoma (Australodrepa) oceanicum oceanicum
Perroud & Montrouzier, 1864
West Australia: Cue Milly Soak, 6.2.2003, Hovorka lg.
Calosoma (Australodrepa) oceanicum oceanicum
Perroud & Montrouzier, 1864
West Australia: Cue Milly Soak, 6.2.2003, Hovorka lg.
Calosoma (Australodrepa) oceanicum oceanicum
Perroud & Montrouzier, 1864
West Australia: Broome, 14.2.91
Calosoma (Australodrepa) oceanicum oceanicum
Perroud & Montrouzier, 1864
West Australia: S. Kimberley, Broome env. , XII.2011
Calosoma (Australodrepa) oceanicum oceanicum
Perroud & Montrouzier, 1864
West Australia: S. Kimberley, Broome env. , XII.2011


Calosoma (Australodrepa) oceanicum klynstrai Breuning, 1927

Calosoma oceanicum klynstrai Breuning, 1927: 158 (type: Tanimbar; coll. Vogt, Zoölogisch Museum, Universiteit van Amsterdam)
Calosoma (Australodrepa) timorensis (= klinstrai) Lapouge, 1932: 407
Australodrepa oceanicum klynstrai Jeannel, 1940: 74

Length 22-28 mm. It has less transverse pronotum and more elongated body and the color of upper body is clearer, costantly green-bronze.
oceanicum klynstrai is found in, South eastern Moluccans islands: Sermata (Hackel et Al, 2005) and in Tanimbar archipelago (Yamdena).

Material examined:
Indonesia: Tanimbar archipelago (Yamdena island)

Notes: Nocturnal, winged. Active individuals were captured in December and later in March, just before the beginning of the raining season.

Calosoma (Australodrepa) oceanicum klynstrai
Breuning, 1927
Indonesia: Tanimbar, Isola Yamdena, XII.08
Calosoma (Australodrepa) oceanicum klynstrai
Breuning, 1927
Indonesia: Tanimbar, Isola Yamdena, III.2009


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