Calosoma (Caminara) denticolle Gebler, 1833

C. denticolle has two tertiary intervals on each sides of a secondary one ("pentaploïde" type). The intervals, that are deeply incised, are all of the same width and height ("homodyname" type), and all of them are transformed into series of rough granules, more or less aligned. C. denticolle can be easily distinguished from the other species collected in Caminara by the acute basal angles of the pronotum, by the body shape stout and convex and by the upper body with noticeable bronzed reflexes with large coppery metallic foveae on the primary intervals, and finally by the male anterior tarsi not dilated.
C. denticolle suensoni, from the Chinese Southern region of Guangdong, should be characterized by the roughness of head and pronotum and by the shape of the pronotum that should be more shorter and wider, narrowed to the base with more pointed posterior angles. These characters are very variable in the specimens of the entire area of diffusion and the elevation of the population of Yuncheng to the rank of subspecies appears superfluous.
C. denticolle occupies a large area, from the Baltic sea to Black sea, and from the coasts of the Black sea to Far East trought southern Russia, Caucasus, northern Iran, Central Asia, southern Siberia, Mongolia, North China. The northernmost records consist in some few specimens found along the southern sea coast of Finland, possibly dragged by strong gales in such an unsuitable environment.

Examined specimens and literature’s data
Armenia. Yerevan, (Breuning, 1927: 229)
Belarus. Turov env., Arevithcy vill., Polesie State Radioecological Reserve (Aleksandrowicz, 2011: 478)
Bulgaria. Southern Black Sea coast: Arkutino, Sozopol (Guéorguiev & Guéorguiev, 1995: 41); Nesebar (http://www.ebay.it)
China. Guangdong: Yüncheng (paratype of denticolle suensoni) (NMB); Hebei (Häckel, 2017: 66); Heilongjiang: Mt.Old father, Mudanjiang (SB); Inner Mongolia (Häckel, 2017: 66); Liaoning: Xiuyan (SB); Ningxia: Dawukou (Deuve, 1997: 56); Shanxi (Häckel, 2017: 66); Sichuan: Hanyuan (Ebay, 2014); Xinjiang: Borochoro chain (Breuning, 1927: 229)
Finland. Tvärminne, Brännskär (Lindroth & Bangsholt, 1985: 49)
Georgia. Abkhazia, (Breuning, 1927: 229)
Iran. Golestan: Gonbad (Ghahari et Al 2009: 439)
Kazakhstan. Aksaj Karačaganak (SB); Dzanybek (SB); Semipalatinsk (=Semej) (SB); Almaty: Kashengel (SB), Aydarly (SB), Panfilov distr. (SB); Aktobe (www.inaturalist.org); Astana (www.inaturalist.org)
Moldova. Southern Moldova (Neculiseanu et Al, 2000: 40)
Mongolia. 50 km N Ulaanbaatar (= Ulan Bator) (SB); 25 km. NE Tsejserteg, Arkhangay (SB); 100 km. SSW Baruun-Urt (SB), Domogov reg., 2km SE Khuvsgol (SB)
Romania. Dobrogea: Tuzla (Jeannel, 1940: 120), Mangalia (SB)
Russia. southern Russia (EM); Altai: Biysk distr. (SB), Aleisk (SB), Loktevsky distr (Gebler, 1833: 274), Olkhovskiy distr (http://molbiol.ru/); Astrachan reg: 25 Km NW Kharabali (SB),70 km S. Astrachan' (SB); Bashkortostan: Ufa (www.inaturalist.org), Tuymazinsky distr. (Instagram, Burns-Scarabaeinae); Bouriazia: Buren river (near the Mongolian border) (SB), Kiakhta (Jeannel, 1940: 120); Čeljabinsk reg: Kizel'skij distr. (www.gbif.org), Čeljabinsk distr. (www.inaturalist.org); Dagestan (Breuning, 1927: 229): Gergebil'skij distr. (www.gbif.org); Irkutsk: Zalari, Irkutsk, Bolshoe Golustnoe (www.inaturalist.org/); Kalmykia: Khulkluta vill. (SB), Ulan-Khol lake (SB); Krasnodar reg: Krasnoyarsk (www.inaturalist.org), Sotschi (VV), Leningradskaya (SB); Orenburg reg: Akbulak distr (SB); Omsk reg: Bolsheukovsky distr.; Rjasan; Viatka (Breuning, 1927: 229); Saint Petersburg env. (SB); Tomsk (Breuning, 1927: 229); Saratov reg: Saratov distr., Balashov distr., Krasnoarmeyskiy distr., Rtischevsky distr. (Sazhnev, 2007: 351, Balakovskiy distr. (www.inaturalist.org); Tatarstan: Nizhnekamskiy distr. (www.inaturalist.org); Tjumen: Tjumen distr (www.inaturalist.org); Volgograd: Volgograd distr (SB), Olkhovskiy distr. (SB).
Turkey. near Istanbul (Jeannel, 1940: 120); Artvin prov: Hopa (LT)
Turkmenistan. (Kryzhanovsky & Atamuradov, 1994: 416); Ahal prov. (www.gbif.org)
Ucraine. Sevastopol (Jeannel, 1940: 120); Poltawa (Breuning, 1927: 229), Kherson (SB), Dnipropetrovsk (www.inaturalist.org); Kharkiv: Devorichna (www.inaturalist.org).
Uzbekistan. Khorezm reg., Karakapakstan rep. (Khamraev, 2003: 47)

Notes: Winged. Adults are mainly active from May to September (according to the material examined) and they can be captured in pit traps.

Calosoma (Caminara) denticolle ♂ Gebler, 1833 Romania: Mangalia, 14-16.8.90, T. Tračka lgt.

Calosoma (Caminara) denticolle ♀ Gebler, 1833 Kazakhstan: Aksaj Karačaganak, VI.88, Čermàk lgt.

Calosoma (Caminara) denticolle ♀ Gebler, 1833 China: Yüncheng, 12.IX.1929 Suenson lgt. (paratype of Calosoma (Charmosta) denticolle suensoni Mandl, 1981) (coll. Naturhistorischen Museum Basel)